Sing/plural Verb to use with ‘one of + who’

1. He is the only one of those men who is/are  always courteous.

2. He is one of those men who is/are always courteous.

3. Pope Francis is one of the popes who has/have led the Catholic Church for almost two thousand years.

Do we use singular/plural verb in these sentences? ‘Who’ refers to what subject? Please explain.

 

 

 

 

 

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